Olsson, A. A., 1922. The Miocene of Northern Costa Rica with Notes on its General Stratigraphic Relations. Bulletins of American Paleontology, 6 (39        ).
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            New Species-Group Names:
            
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                    bocasensis Olsson, 1922. p. 223 (51). pl. iii. f. 14.
                
 
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                    costaricensis Olsson, 1922. p. 217 (45). pl. iii. f. 3. 9. 
                
                 
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                    harrisi Olsson, 1922. p. 225 (53). pl. iii. f 21. 
                
                 
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                    limonensis Olsson, 1922.  p. 219 (47). pl. iii. f. 7. 
                
                 
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                    musaensis Olsson, 1922. p. 219 (47). pl. i. f. 22. 
                
                 
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                    toroensis Olsson, 1922. p. 220 (48). pl. ii. f. 7. 
                
                 
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                    veatchi Olsson, 1922. p. 216 (44). pl. ii. f. 5.